Markéta Tomková (Popelová)



  • Doctoral Thesis: The relationship between DNA modifications and mutations in cancer, 2017 [PDF]
  • Master Thesis: Software tool for modelling coding and processing of information in auditory cortex of mice, 2013 [PDF]
  • Bachelor Thesis: Knihovna steering technik pro virtuální agenty, 2011 (in Czech) [PDF]

Peer-reviewed publications

The full list can be found here: Google Scholar

Selected manuscripts:


  • Popel M.*, Tomkova M.*, Tomek J.*, Kaiser Ł., Uszkoreit J., Bojar O., Žabokrtský Z. Transforming machine translation: a deep learning system reaches news translation quality comparable to human professionals. Nature communications, 11(1), 1-15, 2020. [PDF]


  • Liu, Y. Siejka-Zielińska P., Velikova G., Bi Y., Yuan F., Tomkova M., ... & Song C. X. Bisulfite-free direct detection of 5-methylcytosine and 5-hydroxymethylcytosine at base resolution. Nature biotechnology, 37(4), 424-429, 2019. [PDF]
  • Popelová J. R., Živná R., Tomková M., Tomek J. Quality of life and depression in adults with repaired complex cyanotic congenital heart disease in the Czech Republic. Cor et Vasa, 61(1), 2019. [PDF]


  • Tomkova M., McClellan M., Kriaucionis S., Schuster-Böckler B. DNA replication and associated repair pathways are involved in the mutagenesis of methylated cytosine. DNA repair, 62, 1-7, 2018. [PDF]
  • Tomkova M., Schuster-Böckler B. DNA modifications: naturally more error prone? Trends in Genetics, 34(8), 627-638, 2018. [HTML]
  • Tomkova M., Tomek J., Kriaucionis S., Schuster-Böckler B. Mutational signature distribution varies with DNA replication timing and strand asymmetry. Genome biology, 19(1), 1-12, 2018. [PDF]


  • Popelová J. R., Tomková M., Tomek J. NT-pro BNP predicts mortality in adults with transposition of the great arteries late after M ustard or S enning correction. Congenital heart disease, 12(4), 448-457, 2017. [PDF]


  • Tomkova M., McClellan M., Kriaucionis S., Schuster-Böckler B. 5-hydroxymethylcytosine marks regions with reduced mutation frequency in human DNA. Elife, 5, e17082, 2016. [PDF]


  • Popelová J. R., Kotaška K., Tomková M., Tomek J. Usefulness of N-terminal pro-brain natriuretic peptide to predict mortality in adults with congenital heart disease. The American journal of cardiology, 116(9), 1425-1430, 2015. [HTML]
  • Tomková M., Tomek, J., Novák, O., Zelenka, O., Syka, J., & Brom, C. Formation and disruption of tonotopy in a large-scale model of the auditory cortex. Journal of computational neuroscience, 39(2), 131-153, 2015. [HTML]


  • Brom C., Babor P., Popelová M., Bída M., Tomek J.: Controlling Three Agents in a Quarrel: Lessons Learnt. In Proceedings of Motion in Games (MIG), Lecture Notes in Computer Science. Volume: 7660, Springer, Heidelberg, pp. 158-169, 2012.
  • Popelová M., Brom C., Tomek J., Bída M.: Toolkit for Teaching Steering Behaviors for 3D Human-like Virtual Agents (Demonstration). In Proceedings of Autonomous Agents and Multiagent Systems (AAMAS), Valencia, Spain, pp. 1477-1478, 2012. [PDF] [POSTER]


  • Popelová M., Bída M., Brom C., Gemrot J., Tomek J.: When a Couple Goes Together: Walk Along Steering. In Proceedings of Motion in Games (MIG), Lecture Notes in Computer Science. Volume: 7060, Springer, Heidelberg, pp. 278-289, ISBN 978-3-642-25089-7, 2011. [PDF]
  • Bída M., Brom C., Popelová M.: StoryFactory – a tool for scripting machinimas in Unreal Engine 2 and UDK. In Proceedings of The Fourth International Conference on Interactive Digital Storytelling (ICIDS), Lecture Notes in Computer Science. Volume: 7069, Springer, Heidelberg, pp. 334-337, ISBN 978-3-642-25288-4, 2011. [PDF]
  • Bída M., Brom C., Popelová M.: To Date or Not to Date? A Minimalist Affect-Modulated Control Architecture for Dating Virtual Characters. In Proceedings of The 11th International Conference on Intelligent Virtual Agents (IVA), Lecture Notes in Computer Science. Volume: 6895, Springer, Heidelberg, pp. 419-425, DOI: 10.1007/978-3-642-23974-8_45, 2011. [PDF]
  • Popelová M., Bída M.: Steering techniky pro virtuální agenty. In: Proceedings of Kognice a umelý život XI. (KUŽ), pp. 207-212, ISBN 978-80-7248-644-1, 2011. [PDF]

Projects from Matfyz

Steering Game and Steering Tool

description: Educational tool for teaching, observing, testing and thereby understanding Steering Behaviors (technique of navigation virtual agents, invented by Craig Reynolds).

participation: I was the main programmer of this tool. The tool was based on a tool, which I developed for my Bachelor Thesis.

durantion: 2011-2012

web site:

Story Factory

description: A tool enabling user to create short animated scenes with virtual characters.

participation: I was one of the programmers of the educational tool Story Factory, author of tutorials (both text and videos). I still participate in teaching courses about this tool (for high-school students or their teachers).

durantion: 2010-2011

web site:

© Markéta Tomková (Popelová) 2009 - 2021